ABSTRACT:
This paper reports on the results of monitoring and mapping changes in the spatial extent of standing surficial water and wetland at Enders and surrounding Lakes, Brown County, Nebraska. The overall purpose of the research was to evaluate the potential of remotely sensed data and Geographic Information Systems (GIS) in providing practical, quick and cost effective solution to the problem of detecting and mapping changes in the areal extent of wetlands caused by flooding within an area of 100sq. miles around Enders Lake over a period of several years.
Among the wetlands are saturated meadows, shallow marshes, and open-water lakes. The wetlands range in size from less than one acre to two thousand acres (Wolfe, 1984). This study is very important because a reduction in the size of wetlands can be of great economic lose to the area ranchers as these wetlands, especially the wet meadows provide abundant and nutritious forage which is used as winter cattle feed. These wetlands also used as grazing sites as well as a source of water to livestock.
The data used in this study were: 1) Landsat Thematic Mapper (TM) images (1986, 1991, and 1992); 2) black-and-white (B&W) aerial photographs (1939, 1954, 1961, and 1968); 3) National Aerial Photography Program, both color-infrared (CIR) and B&W photographs (1989 and 1993); and 4) National Wetland Inventory (NWI) data-sets.
Several standard, commonly used image-analysis techniques were considered. The Tasseled-Cap Transformation (TCT ) was used to identify open surficial water and wetland areas from the TM images and scanned CIR photographs. The transformed images were classified using unsupervised classification. In order to calculate areas of surficial water and wetlands, the classified image was multiplied with another raster image created by rasterizing a vector layer of polygons for each lake.
The study shows that remote sensing data with GIS can be used to successfully identify and measure variations in the extent of open surficial water and wetland areas of (even) very small lakes. The study also shows that wetlands within the study area are dynamic and change frequently. The change in open surficial water has a direct effect on the areal extent of the wetland. For example, in 1993, the areal extent of open surficial water showed a negative change for all the lakes. In the same year the areal extent of wetlands also decreased (regardless of a difference in procedures between digital classification and manual photo interpretation).
Finally, this study also shows that the extent of open surficial water has generally increased over the years (especially in bigger lakes like Enders and Willow). In other smaller lakes, the spatial extent of open water has either increased in most years or stayed the same. Wetland areas, too, either increased or remained the same in most cases. But, wetland areas around the big lakes like Moon lake and Ender lake, have decreased considerably.
I. BACKGROUND:
This document is the final report on the results of a study aimed at monitoring and mapping changes in the spatial extent of standing surficial waters and wetlands at Enders and surrounding Lakes in Brown County, Nebraska. The project was conducted by the Center for Advanced Land Management Information Technology (CALMIT), University of Nebraska-Lincoln for the Sandhills Task Force and the U.S. Fish and Wildlife Service. The purpose of the research was to identify, record, and map changes in the areal extent of surface waters and wetlands within a specific area including Enders Lake and to do so using historical remotely sensed data in conjunction with a Geographic Information System (GIS).
II. METHODOLOGY:
A. Study Area:
The study site is the "Enders marsh" and surrounding area, approximately 100 sq. miles in size, in Brown County, NE (Figure 1). The study site is located in the North-Central portion of the state in the physiographic region known as the Nebraska Sandhills. Specifically, the study area is located between 420 13' 00" and 420 22' 30" N latitude and 1000 00' 00" and 1000 13' 00" W longitude.
Figure 1: Study Area
B. Data Acquisition, Processing, and Analysis
The flow chart, shown as Figure 2, summarizes the steps used in executing the project.
Figure 2: Project Flow Chart
(1) Data Acquisition:
The first phase of the project, to obtain the relevant spectral and spatial data, was begun with an in- house search of the image archives maintained by both CALMIT and the Conservation & Survey Division (CSD), UNL. Three satellite images over the study area, acquired by the Landsat Thematic Mapper (TM) sensor at 30-meters ground resolution, were identified in the CALMIT data archive. These images were acquired by the satellite sensor in 1986, 1991, and 1992 (see Figure 3 for specific dates). A number of black-and-white (B&W) aerial photographs, for the years 1939, 1954, 1961, and 1968 for Brown County, NE, were located in the CSD image archive, although only the 1954 photographs covered most of the study area.
The second step in identifying image resources was to search the holdings of the United States Geological Survey (USGS) EROS Data Center. Two sets of National Aerial Photography Program (both color-infrared (CIR) and B&W) photographs covering the study area for the years 1989 and 1993 were located in that archive and purchased (see Figure 4) . National Wetland Inventory (NWI) data-sets, in vector format, for the study area were also obtained to use as a reference.
(2) Hardware / Software
ERDAS Imagine (ver. 8.3.1) and ESRI's ARC-INFO software on Sun Sparc-20 workstations were used to perform all data analyses.
(3) Pre-Processing of TM Scenes:
Each TM image covers a ground area of approximately 13225 square miles (185 KM X 185 KM or 115 miles X 115 miles) , which is a digital data-set of 350 megabytes. To reduce the processing time and storage space, the TM images were down loaded from compact disk and the study area was "cut" out of
Figure 3: Landsat Thematic Mapper (TM) Images (False Color Composite Show TM Bands 4,3,2).
Figure 4: Image Mosaic of Aerial Photographs (Color Infrared and Black-and-white).
the overall coverage. All TM images were then geo-referenced to Universal Transverse Mercator (UTM) map coordinates.
(4) Pre-Processing of Aerial Color-Infrared and Black-and-White Photographs:
The color-infrared photographs were obtained as 9x9-inch positive film transparencies while the black-and-white photographs were the same size but stored on paper. A total of 25 photographs (either CIR or B&W) was required to cover the entire study area.
The first step, converting the photographs to a digital format, was accomplished by means of a SHARP JX 610 high resolution flatbed color scanner. Each photo was scanned at a resolution of 300 dots per inch (dpi) to get a spatial resolution (on the ground) of 4 meters. Next, all individual photographs were "stitched" together to create a digital mosaic of the whole study area (Figure 4). In order to mosaic photos together, each one had to be geographically referenced to known ground locations and geometrically rectified to a common coordinate system (UTM). Each photo was rectified using 10 to 15 Ground Control Points (GCP's), comprised of man-made features (e.g., road intersections, road junctions, bridges, railroad crossings, etc.) which could clearly be identified on both the image and a reference map (USGS topographic). Creating digital mosaics from individual photographs is labor-intensive and requires large amounts of time. Each photograph was joined to the adjacent photograph in such a way as to preserve the correct positional relationships of the features on each photograph, which is difficult because some distortion is always present near the edges of individual aerial photographs.
(5) Analyses of Digital Satellite and Aircraft Images:
Since, the overall aim of the project was to detect and map changes in the areal extent of standing surficial water and wetlands over time, several standard, commonly used image-analysis techniques were considered. The Tasseled-Cap Transformation (TCT ) is a method to convert an input image acquired by a "multispectral sensor" (i.e.., one operating simultaneously in several spectral regions, such as visible green, visible red, and near-infrared) into an image which has three main output components that highlight "feature classes" such as bare soil, vegetation, and water (Kauth and Thomas 1976, Crist and Kauth 1986, and Crist 1985)
The calculation of TCT, which is a complex procedure involving the linear combination of all the bands of an image, results in six "synthetic output channels" for a Landsat-TM image, where band 1 corresponds to soil brightness, band 2 to vegetation greenness, band 3 to wetness, band 4 to haze in the atmosphere, and bands 5 and 6 to "system noise." Trial and error confirmed, for Brown County, that a combination of first two transformed bands provided a best result for separating vegetation, soil and water. The two transformed bands (synthetic channels) were then used to classify the image subset for the study area into three general classes (soil, vegetation, and water)( Figure 5). These classes were then labeled, evaluated, and verified by comparing them to both black-and-white and color-infrared aerial photographs, and also to NWI results. Areas of standing surficial water and wetlands in and around each lake in the study area were then calculated (Figures 6 and 7). The same procedure was applied for all three TM images (1986, 1991, and 1992).
Figure 5: TM Image Classification.
Figure 6: Open Surficial Water Calculated from TM Images.
Figure 7: Wetland Areas Calculated from TM Images.
The same general procedure was used to calculate open surficial water and wetland areas from the scanned CIR photograph (1989) as was used for the TM images, although the actual calculation of the TCT varied slightly. However, the procedure for the B&W photographs was quite different because they summarize reflectance/radiance of earth surface in only the visible portion of electromagnetic spectrum; i.e., they are not a multispectral product. Therefore, the differences in the representation of water, soil and
vegetation is often minuscule or even non-existent. As a result, it is very difficult to use any digital image-processing technique to process and analyze scanned B&W photographs. In order to accomplish the task of calculating water and wetland areas for 1954 and 1993, a manual image-interpretation method was used. This method involved visual interpretation of ground features by the image analyst and subsequent screen-digitizing of the standing water and emergent-wetland areas. Then, the acreage calculations were done (Figures 8 and 9).
RESULTS AND DISCUSSION:
The area of open surficial water calculated for each lake and for various dates is shown in Table 1(a). There are nineteen lakes in the study area, with six having no name provided on the corresponding USGS map. Therefore, these are simply listed as "noname" in the table. The noname lakes tend to be small (sizes range from less than five acres to around ten acres, except noname #1 which is several hundred acres in size).
Table 1(b) summarizes changes in the areal extent of the open surficial water from one year to the next, as does Figure 10. We found six lakes that have a range in open surficial water area between 250 acres and 700 acres, and another eight with variable areal extents between 50 acres and 200 acres. The remaining five lakes range between 5 and 50 acres in sizes, for the years analyzed.
Results of the calculation of spatial extent of open surficial water show that there is a negative change in the total acreage of thirteen lakes and there is apositive change in the total acreage of four lakes (Crystal, Philbrick, Enders, and Chain), whereas, Rat and Clear lakes remained almost unchanged between 1954 and 1986. The biggest decrease in the open water between 1954 and 1986 occurred in Noname #1 lake which
Figure 8: Open Surficial Water Calculated from Aerial Photographs
Figure 9: Wetland Areas Calculated from Aerial Photographs.
Figure 10: Variations in Total Lake Areas and Open Surficial Water Over the Study Period.
Table1: (a) Area of Open Surficial Water
| No. | Lake_Names | Area (acres) | Area (acres) | Area (acres) | Area (acres) | Area (acres) | Area (acres) |
| B&W(7-21-54) | TM(7-16-86) | CIR(6-15-89) | TM(7-14-91) | TM(4-27-92) | B&W(7-29-93) | ||
| 1 | noname 1 | 467.69 | 368.95 | 387.59 | 366.95 | 305.79 | 176.09 |
| 2 | Moon | 617.75 | 572.44 | 589.52 | 593.35 | 722.56 | 576.83 |
| 3 | Clapper | 94.69 | 81.62 | 163.12 | 101.86 | 352.72 | 98.58 |
| 4 | Twin | 178.32 | 144.56 | 144.57 | 147.67 | 148.34 | 154.71 |
| 5 | Rush | 70.62 | 57.16 | 59.04 | 58.49 | 59.82 | 53.76 |
| 6 | Crystal | 36.82 | 48.93 | 87.57 | 51.15 | 93.41 | 37.90 |
| 7 | Philbrick | 48.29 | 114.09 | 137.06 | 135.22 | 207.72 | 60.88 |
| 8 | Skull | 109.56 | 82.06 | 81.29 | 97.41 | 96.30 | 71.68 |
| 9 | Long | 299.99 | 247.75 | 275.27 | 269.99 | 375.40 | 249.46 |
| 10 | Rat | 443.87 | 443.46 | 457.30 | 293.34 | 495.27 | 497.41 |
| 11 | noname 2 | 0.00 | 12.01 | 8.35 | 14.01 | 16.46 | 18.07 |
| 12 | Enders | 401.17 | 452.35 | 437.79 | 464.36 | 496.83 | 488.91 |
| 13 | Clear | 90.22 | 90.96 | 97.25 | 108.75 | 116.98 | 89.42 |
| 14 | noname 3 | 0.00 | 12.68 | 13.21 | 11.12 | 17.12 | 14.88 |
| 15 | Chain | 65.56 | 70.72 | 69.94 | 71.39 | 83.18 | 66.49 |
| 16 | Willow | 443.11 | 431.22 | 434.33 | 457.24 | 465.70 | 433.72 |
| 17 | noname 4 | 0.00 | 2.22 | 20.53 | 45.37 | 52.26 | 17.43 |
| 18 | noname 5 | 0.00 | 6.23 | 9.40 | 9.56 | 14.01 | 0.00 |
| 19 | noname 6 | 0.00 | 5.78 | 18.09 | 12.01 | 28.02 | 0.00 |
(b) Change in Area of Open Surficial Water
is around 98.74 acres and the biggest increase of around 65.80 acres occurred in Philbrick lake. On the
other hand, most of the lake areas have changed positively during 1986 and 1992. In 1993, most lakes
show a reduction in the extent of open surficial water as compared to previous year. The areal extent of all
lakes increased in 1989 as compared to 1986 except Enders, Chain, and Skull. Enders Lake showed a
reduction of 14.56 acres whereas Chain and Skull lakes shrunk only about three-quarters of an acre.
Between 1989 and 1991, eight lakes out of the total nineteen show a reduction in size, with the biggest
reduction (for Clapper Lake) being about 61 acres. The second-largest reduction occurred at Crystal Lake,
which dropped from 87.57 to 51.15 acres. All the lakes showed an increase in their size during 1991 and
1992 except Noname #1 and Skull Lakes. Noname #1 Lake was reduced 61.16 acres while Skull Lake
showed a reduction of only one acre. A negative change occurred between 1992 and 1993 in almost all
lakes; i.e., there was a reduction in the open surficial water extent. The reason for these changes may be
a difference in the season in which the aerial and satellite images were compiled. For example, the 1992
TM image was acquired in April whereas 1993 image was collected in June. In April (spring season) there
is very little green vegetation standing in wetlands; therefore, the satellites are likely to "see" open water.
On the other hand, in June, the vegetation cover tends to be dense, so a lesser amount of water is detected
by satellite. Another potential source of error was mentioned earlier; the fact that the area calculations for
1993 were the result of manual photo interpretation and screen digitizing. This may also, although doubtful,
cause the reduction shown in the extent of open surficial water between 1992 and 1993. But, if open water
measurements for 1993 are compared with those for 1986, half of the lakes increased in open water extent,
including the big lakes ( e.g., Rat, Enders, Willow, which have areas of more than 250 acres). The rest are
very close to the area in 1986. In summary, although, the areal extent of open surficial water fluctuated Table 2: (a) Area of Wetlands (b) Change in Area of Wetlands among various dates, the overall change is either positive or close to zero. Table 2 summarizes the area measurements of wetlands for the duration of study. For ease of data
analysis, lake names were used to represent wetland areas. The wetland area shows a similar trend as was
shown by open surficial water. The wetland extent was showing an increase among various dates for most
lakes. The area of Clapper marsh, the biggest in the study area, fluctuated between 876.57 acres in 1989
and 1227.18 acres in 1991. Also, this wetland showed maximum variation in area over the years. For
example, in 1954, the total wetland area is 926.41arces and in 1986 it increased to 1105.3 acres, but
decreased to 876.57 acres in 1989. It increased to 1227.l8 acres in 1991, and decreased again to 1158.01
acres in 1992. By 1993, it came further down to 920.36 acres. Again, this big drop in the wetland areas
for 1993 is either due to the differences between results obtained by manual interpretation and digital
classification, or merely the seasonal difference in dates of data acquisition by the aircraft or satellite. Wetland areas around eight lakes show an increase between 1954 and 1993. The biggest increase is
in Clapper lake of 163.95 acres. Out of the remaining seven lakes, four lakes (Noname #1, Moon lake,
Philbrick lake, and Enders lake) showed reduction of wetland area of 468.90 acres, 269.95 acres, 155.41
acres, and 287.98 acres respectively. On the other hand if we compare wetland areas between 1986 and
1993, thirteen lakes shows an increase. Out of the remaining six, five wetlands showed a reduction in the
spatial extent whereas the area of the sixth lake (Noname #1) could not be calculated for 1993 (due to the
fact that there was no visual contrast between the wetland on the photo and bare soil). Also, areas for four
lakes could not be calculated for 1954 either because of unavailability of photograph coverage or due to
the reason explained above. CONCLUSION: Over the years, remote sensing has been used as a tool to map large areas of lakes and wetlands.
Moreover, remote sensing in the form of aerial photography served the purpose of identification, delineation
and measurement of spatial extent of wetlands (e.g., National Wetland Inventory (NWI) ; Reimold et al,
1973). The present study shows that remote sensing data in the form of Landsat-TM along with CIR and
B & W photographs and digital image processing techniques like the Tasseled Cap Transformation and
image classification can be used to successfully identify and measure variations in the extent of open surficial
water and wetland areas of (even) very small lakes. The study also shows that wetlands within the study area are very dynamic and change frequently. The
change in open surficial water has a direct effect on the areal extent of the wetland. For example, in 1993,
the areal extent of open surficial water showed a negative change for all the lakes. In the same year the areal
extent of wetlands also decreased (irregardless of a difference in procedures between digital classification
and manual photo interpretation). Finally, this study also shows that the extent of open surficial water is generally increased over the years
(especially in bigger lakes like Enders and Willow lakes). In other smaller lakes, the spatial extent of open
water has either increased in most years or stayed the same. Wetland areas, too, either increased or
remained the same in most cases. But, wetland areas around the big lakes like Moon lake and Ender lake,
have decreased considerably. REFERENCES: Adeniyi, P. O. 1985. Digital Analysis of Multitemporal Landsat Data for Land-use/Land-cover Classification in a Semi-Arid Area of Nigeria. Photogrammetric Engineering and Remote Sensing. 51:1761-1774. Bleed, A., 1990. Groundwater, in An Atlas of Sand Hills(2nd. edition), Edited by A. Bleed and C. Flowerday, University of Nebraska-Lincoln. Carter, V. 1982. Applications of Remote Sensing to Wetlands, in Remote Sensing for Resource Management, edited by C. Johannsen and J. Sanders, Soil Conservation Society of America. Carter, V. 1977. Coastal Wetlands: The Present and Future Role of Remote Sensing. Proceedings of the 11th International Symposium on Remote Sensing of Environment. Environment Research Institute of Michigan, Ann Arbor, Michigan. 1:301-323. Catallo, W. J. 1993. Ecotoxicology and Wetland Ecosystems: Current Understanding and Future Needs. Environmental Toxicology and Chemistry. 12:2209-2224. Crist, E. P. 1985, A TM Tasseled Cap Equivalent Transformation for Reflectance Factor Data.Remote Sensing of Environment 17:301-306. Crist, E. P., R. C. Cicone 1984, Comparisons of the Dimensionality and Features of Simulated Landsat-4 MSS and TM Data. Remote Sensing of Environment. 14:235-246. Frick, A. L. 1984. Detecting Change in a Louisiana Gulf Coast Marsh Using Remote Sensing. Technical Papers of the 50th Annual Meeting of the ASP. American Society of Photogrammetry, Falls Church, Virginia. 2:596-607. Gilmer, D., E. Work, Jr., J. Colwell, and D. Rebel, 1980. Enumeration of Prairie Wetlands with Landsat and Aircraft Data, Photogrammetric Engineering and Remote Sensing. 46:631-634. Howarth, P. J. And G. M. Wickware 1981. Procedures for Change Detection using Landsat Digital Data.
International J. of Remote Sensing. 2:277-291. Jensen, J., 1996. Introduction to Digital Image Processing (2nd. edition). Englewood Cliffs, NJ:Prentice-Hall. Jensen, J.,M. Hodgson, E. Christensen, H. Mackey, Jr., L. Tinney, and R. Sharitz, 1986. Remote Sensing Inland Wetlands: a multi-spectral approach, Photogrammetric Engineering and Remote Sensing.52:87-100. Kauth, R. J., and G. S. Thomas 1976, The Tasseled Cap- A Graphic Description of the Spectral-Temporal Development of Agricultural Crops as seen by Landsat. Proceedings of the Symposium on Machine Processing of Remotely Sensed Data, Purdue University, West Lafayette, IN,PP: 4B41-4B51. Klemas, V., F. Dailber, D. Bartlett, and R. Rogers, 1975. Coastal Zone Classification from Satellite Imagery, Photogrammetric Engineering and Remote Sensing. 41:499-513. LaGrange, T. 1997. Guide to Nebraska's Wetland and Their Conservation Needs. Nebraska Game and Park Commission, Lincoln, NE. Lillesand, T. And Kiefer, 1987. Remote Sensing and Image Interpretation (2nd. edition). New \York:John Wiley. Lo, T. H. C., F. L. Scarpace, and T. M. Lillesand 1984. Multitemporal Analysis of Landsat Data for Agricultural Land Use/Cover Pattern Mapping. Technical Papers of the 50th Annual Meeting of the ASP. American Society of Photogrammetry, Falls Church, Virginia. 1:288-302. Miller, S. 1990. Land Development and Use. In An Atlas of Sand Hills (2nd. edition), edited by A. Bleed and C. Flowerday, University of braska-Lincoln.
Lake_Names
1954 - 1986
1986-1989
1989 - 1991
1991 - 1992
1992 - 1993
1954 -1993 1
noname 1
-98.74
18.64
-20.64
-61.16
-129.71
-291.61 2
Moon
-45.31
17.08
3.83
129.21
-145.73
-40.92 3
Clapper
-13.07
81.50
-61.26
250.86
-254.14
3.89 4
Twin
-33.76
0.01
3.10
0.67
6.38
-23.61 5
Rush
-13.47
1.88
-0.55
1.33
-6.06
-16.86 6
Crystal
12.11
38.64
-36.42
42.26
-55.50
1.08 7
Philbrick
65.80
22.97
-1.84
72.50
-146.83
12.60 8
Skull
-27.50
-0.77
16.12
-1.11
-24.62
-37.88 9
Long
-52.24
27.52
-5.28
105.42
-125.94
-50.53 10
Rat
-0.41
13.84
-163.96
201.93
2.14
53.54 11
noname 2
12.01
-3.66
5.66
2.45
1.61
18.07 12
Enders
51.19
-14.56
26.57
32.47
-7.92
87.75 13
Clear
0.74
6.29
11.50
8.23
-27.56
-0.80 14
noname 3
12.68
0.53
-2.09
6.00
-2.25
14.88 15
Chain
5.17
-0.78
1.45
11.79
-16.68
0.94 16
Willow
-11.88
3.11
22.91
8.45
-31.98
-9.39 17
noname 4
2.22
18.31
24.84
6.89
-34.83
17.43 18
noname 5
6.23
3.17
0.16
4.45
-14.01
0.00 19
noname 6
5.78
12.31
-6.08
16.01
-28.02
0.00
Lake Names
Area(acres)
Area(acres)
Area(acres)
Area(acres)
Area(acres)
Area(acres)
B&W(7-21--54)
TM(7-16-86)
CIR(6-15-89)
TM(7-14-91)
TM(4-27-92)
B&W(7-29-93) 1
noname 1
468.90
321.81
376.45
493.72
522.41
2
Moon
904.74
627.60
571.06
761.48
860.67
634.79 3
Clapper
926.41
1105.30
876.57
1227.18
1158.01
1090.36 4
Twin
46.08
73.39
92.85
178.14
174.58
64.94 5
Rush
24.98
16.23
50.17
83.40
78.95
37.94 6
Crystal
293.57
268.65
271.80
333.37
366.73
293.77 7
Philbrick
593.58
432.34
462.00
515.51
459.47
438.18 8
Skull
41.87
35.14
137.74
75.84
129.43
74.00 9
Long
439.39
534.86
404.14
582.68
700.32
464.00 10
Rat
150.71
112.98
361.33
167.43
211.94
118.76 11
noname 2
0.00
17.35
38.26
19.57
29.36
27.83 12
Enders
499.38
198.47
265.16
209.94
261.76
211.40 13
Clear
115.63
136.77
174.73
218.39
154.34
116.23 14
noname 3
24.36
1.11
19.98
29.80
50.93
10.12 15
Chain
72.66
50.48
44.18
53.82
97.63
77.49 16
Willow
105.45
80.06
97.07
158.79
217.50
88.17 17
noname 4
0.00
42.70
46.22
50.71
82.51
74.00 18
noname 5
0.00
8.67
24.82
27.35
42.26
41.58 19
noname 6
0.00
44.92
29.99
77.84
59.82
39.63
Lake Names
1954 - 1986
1986-1989
1989 - 1991
1991 - 1992
1992 - 1993
1954-1993 1
noname 1
-147.09
54.64
117.27
28.69
-522.41
-468.90 2
Moon
-277.14
-56.54
190.42
99.19
-225.88
-269.95 3
Clapper
178.90
-228.73
350.61
-69.17
-67.65
163.95 4
Twin
27.31
19.46
85.29
-3.56
-109.65
18.85 5
Rush
-8.74
33.94
33.23
-4.45
-41.01
12.96 6
Crystal
-24.92
3.14
61.57
33.36
-72.96
0.20 7
Philbrick
-161.25
29.66
53.51
-56.04
-21.29
-155.41 8
Skull
-6.73
102.60
-61.90
53.60
-55.43
32.13 9
Long
95.47
-130.72
178.54
117.65
-236.32
24.61 10
Rat
-37.73
248.35
-193.90
44.51
-93.18
-31.95 11
noname 2
17.35
20.91
-18.69
9.79
-1.53
27.83 12
Enders
-300.90
66.69
-55.22
51.82
-50.36
-287.98 13
Clear
21.14
37.96
43.66
-64.05
-38.12
0.60 14
noname 3
-23.25
18.87
9.82
21.13
-40.81
-14.24 15
Chain
-22.18
-6.30
9.64
43.81
-20.14
4.83 16
Willow
-25.39
17.01
61.72
58.71
-129.33
-17.28 17
noname 4
42.70
3.52
4.49
31.80
-8.51
74.00 18
noname 5
8.67
16.15
2.53
14.90
-0.68
41.58 19
noname 6
44.92
-14.93
47.85
-18.01
-20.19
39.63
ACKNOWLEDGMENT:
The author thanks Dr Donald Rundquist for his supervision in finishing this
paper. The author also thanks David Derry and Richard Perk of the Center for Advanced Land Management Information Technologies, Conservation and Survey Division, University of Nebraska-Lincoln for their help in this study. The author acknowledges the funding support of the U.S. Fish and Wildlife Service for this research.
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